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Disordered regions in the IRE1α ER lumenal domain mediate its stress-induced clustering

The result's identifiers

  • Result code in IS VaVaI

    <a href="https://www.isvavai.cz/riv?ss=detail&h=RIV%2F00216224%3A90242%2F24%3A00139153" target="_blank" >RIV/00216224:90242/24:00139153 - isvavai.cz</a>

  • Result on the web

    <a href="https://www.embopress.org/doi/full/10.1038/s44318-024-00207-0" target="_blank" >https://www.embopress.org/doi/full/10.1038/s44318-024-00207-0</a>

  • DOI - Digital Object Identifier

    <a href="http://dx.doi.org/10.1038/s44318-024-00207-0" target="_blank" >10.1038/s44318-024-00207-0</a>

Alternative languages

  • Result language

    angličtina

  • Original language name

    Disordered regions in the IRE1α ER lumenal domain mediate its stress-induced clustering

  • Original language description

    Conserved signaling cascades monitor protein-folding homeostasis to ensure proper cellular function. One of the evolutionary conserved key players is IRE1, which maintains endoplasmic reticulum (ER) homeostasis through the unfolded protein response (UPR). Upon accumulation of misfolded proteins in the ER, IRE1 forms clusters on the ER membrane to initiate UPR signaling. What regulates IRE1 cluster formation is not fully understood. Here, we show that the ER lumenal domain (LD) of human IRE1 alpha forms biomolecular condensates in vitro. IRE1 alpha LD condensates were stabilized both by binding to unfolded polypeptides as well as by tethering to model membranes, suggesting their role in assembling IRE1 alpha into signaling-competent stable clusters. Molecular dynamics simulations indicated that weak multivalent interactions drive IRE1 alpha LD clustering. Mutagenesis experiments identified disordered regions in IRE1 alpha LD to control its clustering in vitro and in cells. Importantly, dysregulated clustering of IRE1 alpha mutants led to defects in IRE1 alpha signaling. Our results revealed that disordered regions in IRE1 alpha LD control its clustering and suggest their role as a common strategy in regulating protein assembly on membranes.

  • Czech name

  • Czech description

Classification

  • Type

    J<sub>imp</sub> - Article in a specialist periodical, which is included in the Web of Science database

  • CEP classification

  • OECD FORD branch

    10608 - Biochemistry and molecular biology

Result continuities

  • Project

  • Continuities

Others

  • Publication year

    2024

  • Confidentiality

    S - Úplné a pravdivé údaje o projektu nepodléhají ochraně podle zvláštních právních předpisů

Data specific for result type

  • Name of the periodical

    EMBO Journal

  • ISSN

    0261-4189

  • e-ISSN

  • Volume of the periodical

    43

  • Issue of the periodical within the volume

    20

  • Country of publishing house

    US - UNITED STATES

  • Number of pages

    31

  • Pages from-to

    4668-4698

  • UT code for WoS article

    001306286100002

  • EID of the result in the Scopus database

    2-s2.0-85203078172