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The cryo-EM structure of Photosystem I from Chromera velia with a bound superoxide dismutase heterodimer

The result's identifiers

  • Result code in IS VaVaI

    <a href="https://www.isvavai.cz/riv?ss=detail&h=RIV%2F60076658%3A12310%2F25%3A43910778" target="_blank" >RIV/60076658:12310/25:43910778 - isvavai.cz</a>

  • Alternative codes found

    RIV/00216208:11310/25:10510835

  • Result on the web

    <a href="https://www.nature.com/articles/s41467-025-67637-0" target="_blank" >https://www.nature.com/articles/s41467-025-67637-0</a>

  • DOI - Digital Object Identifier

    <a href="http://dx.doi.org/10.1038/s41467-025-67637-0" target="_blank" >10.1038/s41467-025-67637-0</a>

Alternative languages

  • Result language

    angličtina

  • Original language name

    The cryo-EM structure of Photosystem I from Chromera velia with a bound superoxide dismutase heterodimer

  • Original language description

    Photosystem I is a key component of the solar energy conversion machinery in oxygenic photosynthesis, and its core, where photochemistry occurs, is highly conserved. However, the coral-associated alga Chromera velia that is evolutionary linked to parasitic apicomplexans, exhibits Photosystem I with unusual features. These include the splitting of the central PsaA subunit and the binding of superoxide dismutases as regular subunits. The organization of such a unique Photosystem I was enigmatic. Here, we present the cryo-EM structure of Chromera velia Photosystem I at 1.84 &amp; Aring; resolution. Our work reveals a superoxide dismutase heterodimer bound to the stromal side of the core, stabilized by extensions of canonical subunits, a novel protein PsaV, and a reduced light-harvesting apparatus. We elucidate how the complex evolved to accommodate the superoxide dismutase, assemble the split PsaA, and integrate antenna proteins in a non-canonical orientation. Based on our data and prior physiological data, we propose that this specialized Photosystem I functions likely as an Mehler machine, redirecting electrons from Photosystem II back to water. This mechanism enables Chromera velia to manage redox imbalance and reduce photorespiration through localized oxygen consumption.

  • Czech name

  • Czech description

Classification

  • Type

    J<sub>imp</sub> - Article in a specialist periodical, which is included in the Web of Science database

  • CEP classification

  • OECD FORD branch

    10606 - Microbiology

Result continuities

  • Project

  • Continuities

    I - Institucionalni podpora na dlouhodoby koncepcni rozvoj vyzkumne organizace

Others

  • Publication year

    2025

  • Confidentiality

    S - Úplné a pravdivé údaje o projektu nepodléhají ochraně podle zvláštních právních předpisů

Data specific for result type

  • Name of the periodical

    Nature Communications

  • ISSN

    2041-1723

  • e-ISSN

    2041-1723

  • Volume of the periodical

    17

  • Issue of the periodical within the volume

    1

  • Country of publishing house

    DE - GERMANY

  • Number of pages

    16

  • Pages from-to

    nestránkováno

  • UT code for WoS article

    001668141300002

  • EID of the result in the Scopus database

    2-s2.0-105028521458