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Functional analysis of methyltransferase involved in SL RNA modification in Trypanosoma brucei

Project goals

The trypanosomatid SL RNA plays an essential role in trans-splicing and processing of all mRNA. SL RNA provides the m7G-capped SL sequence to the 5'end of every mRNA. The cap structure of the SL RNA is unique in eukaryotes with 4 nucleotides after the m7G cap carrying a total of seven methyl groups and by convention is referred to as "cap 4". The RNA methyltransferases that catalyze cap 4 modifications are unique to trypanosomatids. It is likely, that the enzymes involved in cap 4 formation, play a pivotal role in regulating the availability of trans-splicing "competent" SL RNA and, therefore, these enzymes might be major players in regulating trans-splicing activity as a whole. Recently, several cap 4 modifying enzymes have been characterized. Theaimof this project is to analyze the function of potential SL RNA methyltransferase MT420 in Trypanosoma brucei. The specific aims of the project involve investigation of a role of MT420 for SL RNA biogenesis and trans-splicing, determination its

Keywords

T. bruceimethyltranserasesSL RNAtrans-splicingcap structureRNA interference

Public support

  • Provider

    Czech Science Foundation

  • Programme

    Post-graduate (doctorate) grants

  • Call for proposals

    Postdoktorandské granty 8 (SGA02008GA1PD)

  • Main participants

  • Contest type

    VS - Public tender

  • Contract ID

    204/08/P585

Alternative language

  • Project name in Czech

    Funkční analýza methyltransferázy účastnící se modifikace SL RNA u Trypanosoma brucei

  • Annotation in Czech

    SL RNA u trypanosomatid má významnou roli při "trans-sestřihu" a  úpravách mRNA. SL RNA poskytuje každé mRNA tzv. SL sekvenci. SL sekvence je krátký úsek SL RNA charakterizovaný m7G čepičkou na 5´-konci následovanou 4 nukleotidy nesoucími sedm methylových skupin. Tato SL RNA čepička je označována jako "cap 4" a je mezi eukaryotickými organismy jedinečná. RNA methyltransferázy, které katalyzují modifikace SL sekvence u trypanosom jsou rovněž unikátní. Je pravděpodobné, že tyto enzymy, účastnícíse modifikace SL sekvence, hrají hlavní roli v regulaci dostupnosti SL RNA pro "trans-sestřih" a mohou tak být důležitými aktéry v regulaci tohoto zásadního procesu. Během posledních let bylo charakterizováno několik enzymů účastnících se modifikace čepičky SL RNA. Cílem tohoto projektu je analyzovat funkci potenciální methyltransferázy MT420 účastnící se modifikace SL sekvence u Trypanosoma brucei - významného parazitického prvoka savců.

Scientific branches

  • R&D category

    ZV - Basic research

  • CEP classification - main branch

    EB - Genetics and molecular biology

  • CEP - secondary branch

    CE - Biochemistry

  • CEP - another secondary branch

  • 10603 - Genetics and heredity (medical genetics to be 3)
    10604 - Reproductive biology (medical aspects to be 3)
    10605 - Developmental biology
    10608 - Biochemistry and molecular biology
    10609 - Biochemical research methods
    30101 - Human genetics

Completed project evaluation

  • Provider evaluation

    O - Nesplněno zadání, smlouva však byla dodržena

  • Project results evaluation

    This project has focused on the characterization of the putative methyltransferase MT420 in Trypanosoma brucei, a human pathogenic parasite.  MT420 is annotated as a hypothetical protein and was originally thought to play a role in the methylation of theunique cap structure of all T. brucei mRNAs.  However, this hypothesis was shown to be incorrect. Bioinformatics have allowed us to identify hig

Solution timeline

  • Realization period - beginning

    Jan 1, 2008

  • Realization period - end

    Dec 31, 2010

  • Project status

    U - Finished project

  • Latest support payment

    Apr 16, 2010

Data delivery to CEP

  • Confidentiality

    S - Úplné a pravdivé údaje o projektu nepodléhají ochraně podle zvláštních právních předpisů

  • Data delivery code

    CEP11-GA0-GP-U/04:3

  • Data delivery date

    Mar 20, 2015

Finance

  • Total approved costs

    1,035 thou. CZK

  • Public financial support

    1,035 thou. CZK

  • Other public sources

    0 thou. CZK

  • Non public and foreign sources

    0 thou. CZK

Basic information

Recognised costs

1 035 CZK thou.

Public support

1 035 CZK thou.

100%


Provider

Czech Science Foundation

CEP

EB - Genetics and molecular biology

Solution period

01. 01. 2008 - 31. 12. 2010